I think Nev might be onto it. There is some tiny tiny mite or something on the underside of the leaves. Dust particle-sized. This is the best magnification I could get with my camera:
The plants are still cropping far more than I can deal with (I'm finding it hard to get excited about
another meal with courgettes in it, just now), and it's not affecting the whole plant by any means.
More concerning is what I found on the underside of the parsnip leaves, which seems to be a different type of mite, and seems to be swiftly killing off the green stuff. This is a worry, because these plants may have to be in the ground for another six months yet. This pic is similarly magnified:
Any ideas about this one?